Publications

The Leatherback Trust’s approach to conservation grounded in scientific research is demonstrated in a vast array of books, journal articles, scholarly texts and management reports.

Books

  • Spotila, J. R. 2004. Sea Turtles: A Complete Guide to Their Biology, Behavior and Conservation. Johns Hopkins University Press, Baltimore, MD. 228pp.

  • Spotila, J. R. 2011. Saving Sea Turtles: Extraordinary Stories from the Battle Against Extinction. Johns Hopkins University Press, Baltimore, MD. 216pp.

  • Spotila, J.R. and Santidrián Tomillo, P. (eds) 2015. Biology and Conservation of Leatherback Turtles. 2015. Johns Hopkins University Press, Baltimore, MD. 228pp.

Articles

2017

  • Hill, J. E., N. J. Robinson, C. M. King, F. V. Paladino. 2017. Diving Behavior and Thermal Habitats of Gravid Hawksbill Turtles at St. Croix, USA. Marine Biology 164:17.

  • Robinson, N. J., F. V. Paladino, and P. Santidrián Tomillo. 2017. Evidence of a Green Turtle Starting its Post-nesting Migration Without Laying all its Vitellogenic Follicles. Marine Turtle Newsletter 152:8-10.

  • Robinson, N. J., K. R. Stewart, P. H. Dutton, R. Nel, F. V. Paladino, and P. Santidrián Tomillo. 2017. Standardising Curved Carapace Length Measurements for Leatherback Turtles, Dermochelys coriacea, to Investigate Global Patterns in Body Size. Herpetological Journal 26:133–136.

2016

  • Robinson, N. J., E. A. Lazo-Wasem, F. V. Paladino, J. D. Zardus, and T. Pinou. 2016. Assortative epibiosis of leatherback, olive ridley and green sea turtles in the Eastern Tropical Pacific. Journal of the Marine Biological Association of the United Kingdom:1-8.

  • Robinson, N. J., R. Majewska, E. A. Lazo-Wasem, R. Nel, F. V. Paladino, L. Rojas, J. D. Zardus, and T. Pinou. 2016. Epibiotic Diatoms Are Universally Present on All Sea Turtle Species. PLoS ONE 11(6).

  • Robinson, N.J., S. J. Morreale, R. Nel, and F. V. Paladino. 2016. Coastal Leatherback Turtles Reveal Conservation Hotspot. Scientific Reports 6:37851.

2015

  • Cyrus, A. Z., J. Swiggs, P. Santidrian Tomillo, F. V. Paladino, and W. S. Peters. 2015. Cannibalism Causes Size-dependent Intraspecific Pressure but Does not Trigger Autotomy in the Intertidal Gastropod Agaronia propatula. Journal of Molluscan Studies 7:1-9.

  • Dornfeld, T. C., N. J. Robinson, P. Santidrian Tomillo, and F. V. Paladino. 2015. Ecology of solitary nesting olive ridley sea turtles at Playa Grande, Costa Rica. Marine Biology 162:123-129.

  • Hill, J. E., F. V. Paladino, J. R. Spotila, and P. Santidrian Tomillo. 2015. Shading and Watering as a Tool to Mitigate the Impacts of Climate Change in Sea Turtle Nests. PLOS ONE.

  • Neeman, N., J. R. Spotila, and M. P. O'Connor. 2015. A Simple, Physiologically-based Model of Sea Turtle Remigration Intervals and Nesting Population Dynamics: Effects of Temperature. Journal of Theoretical Biology 380:516-523.

  • Neeman, N., N. J. Robinson, F. V. Paladino, J. R. Spotila, and M. P. O’Connor. 2015. Phenology Shifts in Leatherback Turtles (Dermochelys coriacea) Due to Changes in Sea Surface Temperature. Journal of Experimental Marine Biology and Ecology.

  • Rivas, M., P. Santidrian Tomillo, J. Dieguez-Uribeondo, and A. Marco. 2015. Leatherback Hatchling Sea-finding in Response to Artificial Lighting: Interaction Between Wavelength and Moonlight. Journal of Experimental Marine Biology and Ecology 463:143-149.

  • Robinson, N. J. and C. Figgener. 2015. Plastic Straw Found Inside the Nostril of an Olive Ridley Sea Turtle. Marine Turtle Newsletter 147:4-6.

  • Santidrian Tomillo, P., M. Genovart, F. V. Paladino, J. R. Spotila, and D. Oro. 2015. Climate Change Overruns Temperature Resilience in Sea Turtles and Threatens Their Survival. Global Change Biology 21:2980-2988.

  • Santidrian Tomillo, P., S. A. Roberts, R. Hernandez , J. R. Spotila, and F. V. Paladino. 2015. Nesting Ecology of East Pacific Green Turtles at Playa Cabuyal, Gulf of Papagayo, Costa Rica. Marine Ecology 36:506-516.

2014

  • Cyrus, A. Z., and W. S. Peters. 2014. The cannibalistic snail Agaronia propatula (Caenogastropoda, Mollusca) is reluctant to feed on autotomized ‘tails’ of conspecifics. Marine and Freshwater Behaviour and Physiology 45(5):285-290.

  • Hill, J. E. 2014. Internesting Diving Behavior and Population Structure of Hawksbill Sea Turtles (Eretmochelys imbricata) on St. Croix, USVI. MS Thesis. Purdue University, Fort Wayne, IN.

  • Keene, E. L., T. Soule, and F. V. Paladino. 2014. Microbial isolations from olive ridley (Lepidochelys olivacea) and East Pacific green (Chelonia mydas agassizii) sea turtle nests in Pacific Costa Rica, and testing of cloacal fluid antimicrobial properties. Chelonian Conservation and Biology 13 (1): 49-5.

  • Robinson, N. J., S. E. Valentine, P. Santidrián Tomillo, V. S. Saba, J. R. Spotila, and F. V. Paladino. 2014. Multidecadal trends in the nesting phenology of Pacific and Atlantic leatherback turtles are associated with population demography. Endangered Species Research. 24:197-206.

  • Santidrián Tomillo, P., D. Oro, F. V. Paladino, R. Piedra, A. E. Sieg, and J. R. Spotila. 2014. High beach temperatures increase female-biased primary sex ratios but reduce output of female hatchlings in the leatherback turtle. Biological Conservation 176:71-79.

  • Santidrián Tomillo, P., S. A. Roberts, R. Hernandez, J. R. Spotila, and F. V. Paladino. 2014. Nesting ecology of east Pacific green turtles at Playa Cabuyal, Gulf of Papagayo, Costa Rica. Marine Ecology.

2013

  • Backof, T. F. 2013. Tracking and Analysis of the Spatial and Thermal Habitats of Inter-nesting Loggerheads (Caretta caretta) in Kyparissia Bay, Greece. Masters Thesis. Purdue University, Fort Wayne, IN.

  • Bryan, J. 2013. Concentrations of heavy metals in scute samples from nesting female olive ridley, Lepidochelys olivacea, and eastern Pacific green, Chelonia mydas agassizii, sea turtles in Costa Rica. Masters Thesis. Purdue University, Fort Wayne, IN.

  • Dapp, D., R. Arauz, M. P. O’Connor, and J. R. Spotila. 2013. Impact of Costa Rican longline fishery on its bycatch of sharks, stingrays, bony fish and olive ridley turtles. Journal of Experimental Marine Biology and Ecology. 448: 228–39.

  • Roe, J. H., P. R. Clune, and F. P. Paladino. 2013. Characteristics of a leatherback nesting beach and implications for coastal development. Conservation and Biology 12(1):34-43.

  • Suss, J. S. 2013. Biophysical and physiological ecology of loggerhead turtle nests at Zakynthos and Kyparissia, Greece. Ph.D. Dissertation. Drexel University, Philadelphia, PA.

2012

  • Blanco, G. S., S. J. Morreale, H. Bailey, J. A. Seminoff, F. V. Paladino, and J. R. Spotila. 2012. Post-nesting movements and feeding grounds of a resident East Pacific green turtle, Chelonia mydas, population from Costa Rica. Endangered Species Research 18: 233-245.

  • Blanco, G. S., S. J. Morreale, J. A. Seminoff, F. V. Paladino, R. A. Piedra, and J. R. Spotila. 2012. Movements and diving behavior of internesting green turtles along Pacific Costa Rica. Integrative Zoology. 8(3): 293-306.

  • Cyrus, A. Z., S. D. Rupert, A. S. Silva, M. Graf, J. C. Rappaport, F. V. Paladino, and W. S. Peters. 2012. The behavioural and sensory ecology of Agaronia propatula (Caenogastropoda: Olividae), a swash-surfing predator on sandy beaches of the panamic faunal province. Journal of Molluscan Studies 78(3):235-245.

  • Keene, E. L. 2012. Microorganisms from sand, cloacal fluid and eggs of Lepidochelys olivacea and standard testing of cloacal fluid antimicrobial properties. MS Thesis. Purdue University, Fort Wayne, IN.

  • Saba, V. S., C. A. Stock, F. V. Paladino, J. R. Spotila, and P. Santidrián Tomillo. 2012. Projected response of an endangered marine turtle population to climate change. Nature Climate Change 2: 814-820.

  • Santidrián Tomillo, P., V. S. Saba, G. S. Blanco, C. A. Stock, F. V. Paladino, and J. R. Spotila. 2012. Climate driven egg and hatchling mortality threatens survival of Eastern Pacific leatherback turtles. PLoS ONE 7(5): e37602.

  • Troost, A. I., S. D. Rupert, A. Z. Cyrus, B. F. Dattilo, and W. S. Peters. 2012. What can we learn from confusing Olivella columellaris and O. semistriata (Olivellidea, Gastropoda), two key species in panamic sandy beach ecosystems?. Biota Neotropica 12(2): 101–113.

  • Valverde, R. A., C. M. Orrego, M. T. Tordoir, F. M. Gómez, D. S. Solís, R. A. Hernández, G. B. Gómez, L. S. Brenes, J. P. Baltodano, L. G. Fonseca, and J. R. Spotila. 2012. Olive ridley mass nesting ecology and egg harvest at Ostional beach, Costa Rica. Chelonian Conservation and Biology 11:1-11.

2011

  • Blanco, G. S., S. J. Morreale, E. Vélez, R. Piedra, W. M. Montes, F. V. Paladino, and J. R. Spotila. 2011. Reproductive output and ultrasonography of an endangered population of East Pacific green turtles. The Journal of Wildlife Management 76: 841–846.

  • Dornfeld, T. C. 2011. Nesting ecology of solitary olive ridleys, Lepidochelys olivacea, within Parque Nacional Marino Las Baulas. MS Thesis. Purdue University, Fort Wayne, IN.

  • Honarvar, S., J. R. Spotila, and M. P. O'Connor. 2011. Microbial community structure in sand on two olive ridley arribada nesting beaches: Playa La Flor, Nicaragua and Playa Nancite, Costa Rica. Journal of Experimental Marine Biology and Ecology 409: 339-344.

  • Rafferty, A. R., P. Santidrián Tomillo, J. R. Spotila, F. V. Paladino, and R. D. Reina. 2011. Embryonic death is linked to maternal identity in the leatherback turtle (Dermochelys coriacea). PLoS ONE 6(6): e21038.

  • Roe, J. H., N. S. Sill, M. R. Columbia, and F. V. Paladino. 2011. Trace metals in eggs and hatchlings of Pacific leatherback turtles (Dermochelys coriacea) nesting at Playa Grande, Costa Rica. Chelonian Conservation Biology 10: 3-9.

  • Rupert, S. D., and W. S. Peters. 2011. Autotomy of the posterior foot in Agaronia (Caenogastropoda: Olividae) occurs in animals that are fully withdrawn into their shells. Journal of Molluscan Studies 77(4):437-440.

  • Sieg, A. E., C. A. Binckley, B. P. Wallace, P. Santidrián Tomillo, R. D. Reina, F. V. Paladino, and J. R. Spotila. 2011. Sex ratios of leatherback turtles: hatchery translocation decreases metabolic heating and female-bias. Endangered Species Research 15:195-204.

  • Soslau, G. R., J. Russell, J. R. Spotila, A. Mathew, and P. Bagsiyao. 2011. Acinetobacter sp. HM746599 isolated from leatherback turtle blood. FEMS Microbiology Letters. 322 (2): 166-171.

  • Soslau, G., J. R. Spotila, A. Chun, S. Yi, and K. T. Weber. 2011. Potentially lethal bacteria in leatherback turtle eggs in the wild threaten both turtles and conservationists. Journal of Experimental Marine Biology and Ecology 410: 101–106.

2010

  • Blanco, G. S. 2010. Movements and behavior of the East Pacific green turtle (Chelonia mydas) from Costa Rica. Ph.D. Dissertation. Drexel University, Philadelphia, PA.

  • Garrett, K., B. P. Wallace, J. Garner, and F. V. Paladino. 2010. Variations in leatherback turtle nest environments: consequences for hatching success. Endangered Species Research 11: 147-155.

  • Hamann , M., M. H. Godfrey, J. A. Seminoff, K. Arthur, P. C. R. Barata, K. A. Bjorndal, A. B. Bolten, A. C. Broderick, L. M. Campbell, C. Carreras, P. Casale, M. Chaloupka, S. K. F. Chan, M. S. Coyne, L. B. Crowder, C. E. Diez , P. H. Dutton, S. P. Epperly, N. N. FitzSimmons, A. Formia, M. Girondot, G. C. Hays, I. S. Cheng, Y. Kaska, R. Lewison, J. A. Mortimer, W. J. Nichols, R. D. Reina, K. Shanker, J. R. Spotila, J. Tomás, B. P. Wallace, T. M. Work, J. Zbinden, and B. J. Godley. 2010. Global research priorities for sea turtles: informing management and conservation in the 21st century. Endangered Species Research 11: 245-269.

  • Santidrián Tomillo, P., F. V. Paladino, S. J. Suss, and J. R. Spotila. 2010. Predation of leatherback turtle hatchlings on the crawl to the water. Chelonian Conservation Biology 9: 18-25.

  • Sieg, A. E. 2010. Physiological constraints on the ecology of activity-limited ectotherms. Ph.D. Dissertation. Drexel University, Philadelphia, PA.

  • Sieg, A. E., E. Zandona, V. M. Izzo, F. V. Paladino, and J. R. Spotila. 2010. Population level “flipperedness” in the Eastern Pacific leatherback turtle. Behavioural Brain Research 206: 135-138.

  • Valentine, S. E. 2010. Analysis of climate change on leatherback (Dermochelys coriacea) nesting patterns at Parque Nacional Marino Las Baulas, Costa Rica. MS Thesis. Purdue University, Fort Wayne, IN.

2009

  • Fonseca, L. G., G. A. Murillo, L. Guadamuz, R. M. Spinola, and R. A. Valverde. 2009. Downward but stable trend in the abundance of arribada olive ridley sea turtles (Lepidochelys olivacea) at Nancite beach, Costa Rica (1971-2007). Chelonian Conservation and Biology 8(1): 19-27.

  • Reina, R. D., J. R. Spotila, F. V. Paladino, and A. E. Dunham. 2009. Changed reproductive schedule of leatherback turtles, Dermochelys coriacea, in the eastern Pacific following the 1997/98 transition from El Niño to La Niña conditions. Endangered Species Research 7: 155-161.

  • Santidrián Tomillo, P., J. S. Suss, B. P. Wallace, K. D. Magrini, G. S. Blanco, F. V. Paladino, and J. R. Spotila. 2009. Influence of emergence success on the annual reproductive output of leatherback turtles. Marine Biology 156: 2021-2031.

  • Southward, A., and L. Avens. 2009. Physiological, behavioral, and ecological aspects of migration in reptiles. Journal of Comparative Physiology B 180:1–23.

  • Suryan, R. M., V. S. Saba, B. P. Wallace, S. A. Hatch, M. Frederiksen, and S. Wanless. 2009. Environmental forcing on life history strategies: Evidence for multi-trophic level responses at ocean basin scales. Progress in Oceanography 81: 214-222.

  • Wallace, B. P., and V. Saba. 2009. Environmental and anthropogenic impacts on intra-specific variation in leatherback turtles: opportunities for targeted research and conservation. Endangered Species Research 7: 11-21.

2008

  • Garrett, K. 2008. Intra-beach variation in leatherback turtle nest environments: implications for hatchling success and egg relocation strategies. MS Thesis. Indiana Purdue University, Fort Wayne IN.

  • Honarvar, S. 2008. Effects of density on hatching success of olive ridley nests on arribada beaches. Ph.D. Dissertation. Drexel University, Philadelphia, PA.

  • Honarvar, S., M. P. O’Connor, and J. R. Spotila. 2008. Density-dependent effects on hatching success of the olive ridley turtle, Lepidochelys olivacea. Oecologia: 157:221-230. 

  • Kowalczyk, N. 2008. Sensitivity of leatherback, Demochelys coriacea, nesting activity to lunar and tidal cycles. Honours Thesis. Monash University, Melbourne, Australia.

  • Saba, V. S., J. R. Spotila, F. P. Chavez, and J. A. Musick. 2008. Bottom-up and climatic forcing on the worldwide population of leatherback turtles. Ecology: 89:1414-1427.

  • Santidrián Tomillo, P. 2008. Nesting ecology of leatherback turtles at Playa Grande, Costa Rica. Ph.D. Dissertation. Drexel University, Philadelphia, PA.

  • Santidrián Tomillo, P., V. S. Saba, R. Piedra, F. V. Paladino, and J. R. Spotila. 2008. Effects of illegal harvest of eggs on the population decline of leatherback turtles in Las Baulas Marine National Park, Costa Rica. Conservation Biology 22: 1216-1224.

2007

  • Clusella Trullas, S., and F. V. Paladino. 2007. Micro-environment of olive ridley turtle nests deposited during an aggregated nesting event. Journal of Zoology 272:367-376.

  • Piedra, R., E. Vélez, P. Dutton, E. Possardy, and C. Padilla. 2007. Nesting of the leatherback turtle (Dermochelys coriacea) from 1999-2000 through 2003-2004 at Playa Langosta, Parque Nacional Marino Las Baulas de Guanacaste, Costa Rica. Chelonian Conservation and Biology 6: 111-116.

  • Price, E. R., F. V. Paladino, K. P. Strohl, P. Santidrián Tomillo, K. Klann, and J. R. Spotila. 2007. Respiration in neonate sea turtles. Comparative Biochemistry and Physiology A 146: 422-428.

  • Saba, V. S. 2007. Bottom-up and climatic forcing on the nesting and foraging ecology of leatherback turtles (Dermochelys coriacea). Ph.D. Dissertation. Virginia Institute of Marine Science.

  • Saba, V. S., P. Santidrián Tomillo, R. D. Reina, J. R. Spotila, J. A. Musick, D. A. Evans, and F. V. Paladino. 2007. The effect of El Niño Southern Oscillation on the reproductive frequency of eastern pacific leatherback turtles. Journal of Applied Ecology 44: 395-404. 

  • Santidrián Tomillo, P., E. Velez Carballo, R. D. Reina, R. Piedra Chacon, F. V. Paladino, and J. R. Spotila. 2007. Reassessment of the leatherback turtle (Dermochelys coriacea) population nesting at Parque Nacional Marino Las Baulas, Costa Rica: Effects of conservation efforts. Chelonian Conservation and Biology: 6:54-62.

  • Wallace, B. P., P. R. Sotherland, P. Santidrián Tomillo, R. D. Reina, J. R. Spotila, and F. V. Paladino. 2007. Maternal investment in reproduction and its consequences in leatherback turtles. Oecologia 152: 37-47.

2006

  • Clusella, S., J. R. Spotila, and F. V. Paladiino. 2006. Energetics during hatchling dispersal of the olive ridley turtle, Lepidochelys olivacea, using doubly labeled water. Physiological and Biochemical Ecology. 79: 389-399.

  • Dalton, E. 2006. The leatherback turtle, Demochelys coriacea: A model to test age-related reproductive hypothesis. Honours Thesis. Monash University, Melbourne, Australia.

  • Wallace, B. P., J. A. Seminoff, S. S. Kilham, J. R. Spotila, and P. H. Dutton. 2006. Leatherback turtles as oceanographic indicators: stable isotope analyses reveal a trophic dichotomy between ocean basins. Marine Biology: 149:953-960.

  • Wallace, B. P., P. R. Sotherland, P. Santidrián Tomillo, S. S. Bouchard, R. D. Reina, J. R. Spotila, and F. V. Paladino. 2006. Egg components, egg size, and hatchling size in leatherback turtles. Comparative Biochemistry and Physiology A 145: 524-532.

  • Wallace, B. P., S. S. Kilham, F. V. Paladino, and J. R. Spotila. 2006. Energy budget calculations reveal resource limitation in Eastern Pacific leatherback turtles. Marine Ecology Progress Series 318: 263-270.

2005

  • Clune, P. 2005. Natural and human induced variations on beach dynamic and their effects on leatherback sea turtle conservation. MS Thesis. Purdue University, Fort Wayne, IN.

  • Ralph, C. R., R. D. Reina, B. P. Wallace, P. R. Sotherland, J. R. Spotila, and F. V. Paladino. 2005. Effect of egg location and respiratory gas concentrations on developmental success in nests of the leatherback turtle, Dermochelys coriacea. Australian Journal of Zoology 53: 1-6.

  • Reina, R. D., K. J. Abernathy, G. J. Marshall, and J. R. Spotila. 2005. Respiratory frequency, dive behaviour and social interactions of leatherback turtles, Dermochelys coriacea during the inter-nesting interval. Journal of Experimental Marine Biology and Ecology 316: 1-16.

  • Soslau, G., P. J. Prest, R. Class, R. George, F. V. Paladino, and G. Violetta. 2005. Comparison of sea turtle thrombocyte aggregation to human platelet aggregation in whole blood. Comparative Biochemistry and Physiology B 142: 353-360.

  • Wallace, B. P. 2005. Physiological Ecology of Leatherback Turtles. Ph.D. Dissertation. Drexel University, Philadelphia, PA.

  • Wallace, B. P., C. L. Williams, F. V. Paladino, S. J. Morreale, R. T. Lindstrom, and J. R. Spotila. 2005. Bioenergetics and diving activity of internesting leatherback turtles (Dermochelys coriacea) at Parque Nacional Marino Las Baulas, Costa Rica. Journal of Experimental Biology 208: 3873-3884.

2004

  • Drake, D. L., J. E. Behm, M. A. Hagerty, P. A. Mayor, S. J. Goldenberg, and J. R. Spotila. 2004. Marine turtle nesting activity at Playa Naranjo, Santa Rosa National Park, Costa Rica, for the 1998-1999 season. Chelonian Conservation and Biology 4: 675-678.

  • Nordmoe, E. D., A. E. Sieg, P. R. Sotherland, J. R. Spotila, F. V. Paladino, and R. D. Reina. 2004. Nest site fidelity of leatherback turtles at Playa Grande, Costa Rica. Animal Behaviour. 68: 387-394.

  • Price, E. R., B. P. Wallace, R. D. Reina, J. R. Spotila, F. V. Paladino, R. Piedra, and E. Velez. 2004. Size, growth, and reproductive output of adult female leatherback turtles Dermochelys coriacea. Endangered Species Research 5: 1-8.

  • Sill, N. S. 2004. The effect of heavy metals on hatching success of remigrant and new nesting leatherback females in Costa Rica: Are experienced females better? MS Thesis. Purdue University, Fort Wayne, IN.

  • Soslau, G., B. P. Wallace, C. Vincente, S. J. Goldenberg, T. Tupis, J. R. Spotila, R. George, F. V. Paladino, B. Whitaker, G. Violetta, and R. Piedra. 2004. Comparison of functional aspects of the coagulation cascade in human and sea turtle plasmas. Comparative Biochemistry and Physiology B 138: 399-406.

  • Wallace, B. P., P. R. Sotherland, J. R. Spotila, R. D. Reina, B. F. Franks, and F. V. Paladino. 2004. Biotic and abiotic factors affect the nest environment of embryonic leatherback turtles, Dermochelys coriacea. Physiological and Biochemical Zoology 77: 423-432.

2003

  • Bell, B. A., J. R. Spotila, F. V. Paladino, and R. D. Reina. 2003. Low reproductive success of leatherback turtles, Dermochelys coriacea, is due to high embryonic mortality. Biological Conservation 115: 131-138.

  • Bowen , B. W., A. L. Bass, S. Chow, M. Bostrom, K. A. Bjorndal, A. B. Bolten, T. Okuyama, B. M. Bolker, S. Epperley, E. Lacasella, D. Shaver, M. Dodd, S. R. Hopkins-Murphy, J. A. Musick, M. Swingle, K. Rankin-Baransky, W. Teas, W. N. Wetzel, and P. H. Dutton. 2003. Natal homing in juvenile loggerhead turtles (Caretta caretta). Molecular Ecology. 13 (12): 3797-3808.

2002

  • Crim, J. L., J. R. Spotila, M. P. O'Connor, R. D. Reina, C. J. Williams, and F. V. Paladino. 2002. The leatherback turtle, Dermochelys coriacea, exhibits both polyandry and polygyny. Molecular Ecology 11(10): 2097-2106.

  • Drake, D. L., and J. R. Spotila. 2002. Thermal tolerances and the timing of sea turtle hatchling emergence. Journal of Thermal Biology. 27: 71-81.

  • Plotkin, P. T., and J. R. Spotila. 2002. Post-nesting migrations of loggerhead sea turtles, Caretta caretta, from Georgia, U.S.A.: Conservation implications for a genetically distinct subpopulation. Oryx 36: 396-399.

  • Reina, R. D., P. A. Mayor, J. R. Spotila, and F. V. Paladino. 2002. Nesting ecology of the leatherback turtle, Dermochelys coriacea, at Parque Nacional Marino Las Baulas, Costa Rica 1988-89 to 1999-2000. Copeia 2002: 653-664.

  • Reina, R. D., T. T. Jones, and J. R. Spotila. 2002. Salt and water regulation by the leatherback sea turtle Dermochelys coriacea. Journal of Experimental Biology: 205: 1853-1860.

2001

  • Bell, B. A. 2001. Fertility of leatherback turtles at Playa Grande, Costa Rica. MS Thesis. Drexel University, Philadelphia, PA.

  • Bilinski, J. J., R. D. Reina, J. R. Spotila, and F. V. Paladino. 2001. The effects of nest environment on calcium mobilization by leatherback turtle embryos (Dermochelys coriacea) during development. Comparative Biochemistry and Physiology A 130: 151-162.

  • Drake, D. L., and J. R. Spotila. 2001. Thermal tolerances and the timing of sea turtle hatchling emergence. Journal of Thermal Biology 27: 71-81.

  • Drake, D. L., M. A. Hagetry, J. E. Behm, and S. J. Goldenburg. 2001. Lepidochelys olivacea (olive ridley sea turtle) predation. Herpetological Review 32: 104.

  • Rankin-Baransky, K., C. J. Williams, A. L. Bass, B. W. Bowen, and J. R. Spotila. 2001. Origin of loggerhead turtles stranded in the Northeastern United States as determined by mitochondrial DNA analysis. Journal of Herpetology: 35:638-646.

  • Rostal, D. C., J. S. Grumbles, K. S. Palmer, V. A. Lance, J. R. Spotila, and F. V. Paladino. 2001. Changes in gonadal and adrenal steroid levels in the leatherback sea turtle (Dermochelys coriacea) during the nesting cycle. General and Comparative Endocrinology 122: 139-147.

2000

  • Spotila, J. R., R. D. Reina, A. C. Steyermark, P. T. Plotkin, and F. V. Paladino. 2000. Pacific leatherback turtles face extinction. Nature: 405: 529-530.

1998

  • Binckley, C. A., J. R. Spotila, K. S. Wilson, and F. V. Paladino. 1998. Sex determination and sex ratios of Pacific leatherback turtles, Dermochelys coriacea. Copeia 1998: 291-300.

  • Penick, D. N., J. R. Spotila, M. P. O'Connor, A. C. Steyermark, R. H. George, C. J. Salice, and F. V. Paladino. 1998. Thermal independence of muscle tissue metabolism in the leatherback turtle, Dermochelys coriacea. Comparative Biochemistry and Physiology A 120: 399-403.

1996

  • Chaves, A., G. Serrano, G. Marin, E. Arguedas, A. Jimenez, and J. R. Spotila. 1996. Biology and conservation of leatherback turtles, Dermochelys coriacea, at Playa Langosta, Costa Rica. Chelonian Conservation and Biology 2: 184-189.

  • Morreale, S. J., E. A. Standora, J. R. Spotila, and F. V. Paladino. 1996. Migration corridor for sea turtles. Nature 384: 319-320.

  • Paladino, F. V., J. R. Spotila, M. P. O'Connor, and R. E. Gatten Jr. 1996. Respiratory physiology of adult leatherback turtles (Dermochelys coriacea) while nesting on land. Chelonian Conservation and Biology 2: 223-229.

  • Rostal, D. C., F. V. Paladino, R. M. Patterson, and J. R. Spotila. 1996. Reproductive physiology of nesting leatherback turtles (Dermochelys coriacea) at Las Baulas National Park, Costa Rica. Chelonian Conservation and Biology 2: 230-236.

  • Spotila, J. R., A. E. Dunham, A. J. Leslie, A. C. Steyermark, P. T. Plotkin, and F. V. Paladino. 1996. Worldwide population decline of Dermochelys coriacea: Are leatherback turtles going extinct?. Chelonian Conservation and Biology 2: 209-222.

  • Steyermark, A. C., K. Williams, J. R. Spotila, F. V. Paladino, D. C. Rostal, S. J. Morreale, M. T. Koberg, and R. Arauz. 1996. Nesting leatherback turtles at Las Baulas National Park, Costa Rica. Chelonian Conservation and Biology 2: 173-183.

1990

  • Paladino, F. V., M. P. O’Connor, and J. R. Spotila. 1990. Metabolism of leatherback turtles, gigantothermy, and thermoregulation of dinosaurs. Nature 344: 858-860.

1985

  • Standora, E. A., and J. R. Spotila. 1985. Temperature-dependent sex determination in sea turtles. Copeia (3): 711-722.

1982

  • Morreale, S. J., G. J. Ruiz, J. R. Spotila, and E. A. Standora. 1982. Temperature-dependent sex determination: Current practices threaten conservation of sea turtles. Science, New series 216 (4551):1245-1247.

Reports

  • Arauz, R., and T. Steiner. 2005. Project to protect and manage 25% of Costa Rica’s exclusive economic zone, including the creation and consolidation of the Cocos-Baulas marine Biological Corridor. Pretoma, San Jose, Costa Rica. 45 pp.

  • Cajiao Jiménez, M. V., R. Salazar Cambronero, M. Valverde Soto, I. Naranjo Vargas, and R. Arauz Vargas. 2003. Régimen legal de los recursos marinos y costeros en Costa Rica. Ambio Fundación. 192 pp.

  • Paladino, F. V., and S. J. Morreale. 2001. Sea Turtles. Encyclopedia of Ocean Sciences,18 pp.

  • Plotkin, P. T. 2001. A marine turtle conservation strategy and action plan for the northern Indian Ocean. IUCN/SSC Marine Turtle Specialist Group Publication No. 3. Washington, DC. 33 pp.

  • Robinson, N. J., and F. V. Paladino. 2013. Sea Turtles. Earth systems and environmental sciences, Elsevier. Reference Module.

  • Rojas, L. A., and M. I. Chavarría. 2005. Corredores biológicos de Costa Rica. Sistema Nacional de Áreas de Conservación, Ministerio del Ambiente y Energía, San Jose, Costa Rica. 46 pp plus maps and appendices.

  • Santidrián Tomillo, P. 2008. From the ocean to the beach: widening the limits of conservation. Endangered Species Update. University of Michigan.

  • Santidrian Tomillo, P., V. S. Saba, C. D. Lombard, J. M. Valiulis, N. J. Robinson, F. V. Paladino, J. R. Spotila, C. Fernandez, M. L. Rivas, J. Tucek, R. Nel, and D. Oro. 2015. Global Analysis of the Effect of Local Climate on the Hatchling Output of Leatherback Turtles. Scientific Reports. In Press.

  • Spotila, J. R. 2001. 21st Annual symposium on sea turtle biology and conservation “Coming of Age,” Philadelphia, USA February 24-28, 2001: President’s Report . Marine Turtle Newsletter 93: 15-17.

  • Spotila, J. R., and F. V. Paladino. 2004. Parque Marino Las Baulas: Conservation lessons from a new national park and from 45 years of conservation of sea turtles in Costa Rica. Chapter 15 in Frankie, G. W., A. Matta, and S. B. Vinson (eds), Biodiversity in Costa Rica: Learning the Lessons in a Seasonal Dry Forest. University of California Press. 194–209.

  • Tiffer Sotomayor, R., R. Piedra Chacón, N. Rodriguez Garro, A. Rosales Ruíz, C. A. Díaz Reyes, L. G. Briceño Aragón, A. Fonseca Matarrita, B. Cortés Carrera, and C. Chang. 2004. Plan de acción del plan de manejo del Parque Nacional Marino Las Baulas de Guanacaste 2005-2010. Centro Científico Tropical, San Jose, Costa Rica, 286 pp.

Theses & dissertations

2015

  • Koval, J. 2015. Use of Microsatellites to Compare Solitary vs. Arribada Nesting Olive Ridley Turtles (Lepidochelys olivacea) Along the Eastern Pacific Coast of Costa Rica. Purdue University.

  • Rhodes, A. 2015. Assessing the Effect of Beach Dynamics on Nest Site Selection of Green Turtles (Chelonia mydas) in Playa Cabuyal, Costa Rica. Purdue University.

  • Swiggs, J. 2015. The Effect of Incubation Conditions on Leatherback (Dermochelys coriacea) and Green (Chelonia mydas) Sea Turtle Hatchling Survival. Purdue University.

2014

  • Clyde-Brockway, C. E. 2014. Inter-nesting and Post-nesting Movements and Behavior of East Pacific Green Turtles (Chelonia mydas agassizii) from Playa Cabuyal, Guanacaste, Costa Rica. Purdue University.